Paul C. Johnson


My interest and training in insect ecology and integrated pest management have led me to conduct research on several insects of economic importance. During a post-doc with Dr. Jack E. Coster in the College of Forestry, Stephen F. Austin State University, Nacogdoches, TX, I did research on the dispersal of the southern pine beetle.

After joining the faculty in the Department of Entomology at the University of New Hampshire in 1979, I shifted my research to apple maggot fly dispersal in apple orchards, as well as supervising graduate research on a variety of other apple pests.

In 1980, a project on the relationship of gypsy moth egg hatch to degree-day accumulation conducted with graduate students in my Insect Ecology course led to an interest in the gypsy moth in general and prompted several years of research in cooperation with Dr. R. Marcel Reeves on the impact of ground beetle predators on gypsy moth larvae. This involved the use of the biological tag rubidium and its transfer from larvae fed on Rb-enriched diet to the predators that fed on them. While involved in this research, I continued to refine the gypsy egg hatch model (unpublished data).

Although no longer actively engaged in research, my most recent research activity involved development of a degree-day model of gypsy moth egg hatch incorporating elevation and slope aspect. It was prompted by a series of mesoscale models developed in the southern United States that incorporated my earlier work and which seemed to have some utility in predicting egg hatch over a varied terrain.

Paul Johnson

Associate Professor,
Natural Resources


  • 1975 Ph.D. Insect Ecology, Cornell University
  • 1968 B.S. Biology, Emory & Henry College

Teaching Responsibilities

  • NR 401: Introduction to Natural Resources
  • NR 410: Insects and Society
  • NR 435: Contemporary Conservation Issues and Environmental Awareness
  • NR 506: Forest Entomology
  • NR 713/813: Quantitative Ecology

Selected Publications

Johnson, P.C. and R.M. Reeves. 1995. Incorporation of the biological marker rubidium in gypsy moth, Lymantria dispar (L.) (Lepidoptera: Lymantriidae), and its transfer to the predator Carabus nemoralis Muller (Coleoptera: Carabidae). Environ. Entomol. 24(1): 46-51. 

Andersen, B.C., D.L. Kulhavy, and P.C. Johnson. 1984. Estimating infestation rates of the Nantucket pine tip moth (Lepidoptera: Tortricidae) through sequential sampling. Environ. Entomol. 13:1593-1597.

Johnson, P.C. 1983. Response of adult apple maggot (Diptera: Tephritidae) to Pherocon A.M. traps and red spheres in a non-orchard habitat. J. Econ. Entomol. 76:1279-1284. 

Johnson, P.C., D.P. Mason, S.L. Radke, and K.T. Tracewski. 1983. Gypsy moth, Lymantria dispar (L.) (Lepidoptera: Lymantriidae), egg eclosion: Degree-day accumulation. Environ. Entomol. 12:929-932.

Kulhavy, D.L., and P.C. Johnson. 1983. Southern pine beetle: Annotated bibliography, 1868-1982. Center for Applied Studies. School of Forestry, Stephen F. Austin State University, Nacogdoches, TX. 

Tracewski, K.T., P.C. Johnson, and A.T. Eaton. 1983. Relative densities of predaceous diptera (Cecidomyiidae, Chamaemyiidae, Syrphidae) and their aphid prey in New Hampshire, U.S.A., apple orchards. Prot. Ecol. 6:199-207.

Florence, L.Z., P.C. Johnson, and J.E. Coster. 1982. Behavioral and genetic diversity during dispersal: Analysis of a polymorphic esterase locus in the southern pine beetle, Dendroctonus frontalis Zimm. Environ. Entomol. 11:10141018.

Geer, S.F., J.E. Coster, and P.C. Johnson. 1981. Effect of weather on flight activity of southern pine beetle. J. Ga. Entomol. Soc. 16:272-282.

Bunt, W.D., J.E. Coster, and P.C. Johnson. 1980. Behavior of the southern pine beetle on the bark of host trees during mass attack. Ann. Entomol. Soc. Amer. 73:647-652.

Johnson, P.C., and J.E. Coster. 1980. Seasonal and behavioral chemical affects on dispersion of the southern pine beetle within infestations. In: Proc. Second IUFRO Conf. on Dispersal of Forest Insects: Evaluation, Theory and Management Implications. Sandpoint, ID. (August 1979). A.A. Berryman and L. Safranyik, eds. pp. 173-193. 

Reeve, R.J., J.E. Coster, and P.C. Johnson. 1980. Spatial distribution of flying southern pine beetle (Coleoptera: Scolytidae) and the predator Thanasimus dubius (Coleoptera: Cleridae). Environ. Entomol. 9:113-118.

Coster, J.E., and P.C. Johnson. 1979. Characterizing flight aggregation of the southern pine beetle. Environ. Entomol. 8:381- 387.

Coster, J.E., and P.C. Johnson. 1979. Dispersion patterns of Dendroctonus frontalis and its predator Thanasimus dubius: Influence of behavioral chemicals. IUFRO Confr. on Dispersal of Forest Insects: Evaluation, Theory and Management Implications. Zurich/Zuoz, Switzerland. (September 1978). In: Bull. Swiss Entomol. Soc. 52:309-322. V. Delucchi and W. Baltensweiler, eds.

Johnson, P.C., and J.E. Coster. 1978. Probability of attack by southern pine beetle in relation to distance from an attractive host tree. For. Sci. 24:574-580. 

Johnson, P.C. 1977. Sex-ratio estimation, sequential sampling, and the programmable pocket calculator. Bull. Entomol. Soc. Amer. 23:251-254.


 Ant Wars!
This middle school or high school level laboratory provides an introduction to scientific method of observation and data analysis using interactions between species and colonies of ants. Materials are readily available without expense (with the exception of several disecting scopes that are useful, but not essential).

Life in Pine Cone
This middle school or high school level laboratory may be easily adapted to the elementary level. It studies the biodiversity within the pine cone community and may serve as an interesting introduction to biodiversity issues on a broader scale. Equipment may be constructed by the student from readily available household materials (with the exception of several dissecting scopes that are necessary for careful examination of the material).

This upper elementary/middle school project demonstrates the life history of an insect, the large milkweed bug, that goes through incomplete (gradual) metamorphosis from egg to adult in about 35 days. Students build a home for their bug, observe it, sketch it, record its length and stages of growth and finally compute the growth rate of a population of milkweed bugs under similar environmental conditions.

 Project Monarch
Project Monarch began as an exchange of projects between elementary schools in Nacogdoches, TX and Dover, NH in 1990 in support of the move to make the Monarch butterfly our national insect. It grew into a nationwide project involving student contributions to a poster that grew to over a half mile in length! Part of the poster was displayed in the Smithsonian Institute Museum of Natural History, Washington, DC, in January 1995.

 NH Ag in the Classroom
As COLSA representative to NH Agriculture in the Classroom, I am involved in their efforts to bring ag literacy resource materials into the classroom.